This phylogeny is the author's opinion based on his interpretation of the literature. It is however, broadly congruent with recent works (e.g. Norell et al., 2001; Rauhut, 2000; Norell and Clarke, 2001; Chiappe, 2001, Holtz et al., 2004; Tykoski, 2005; Carrano and Sampson, 2007; Smith et al., 2007; Brusatte and Sereno, 2008; Benson, 2010). The relationships of modern birds are based on molecular studies.


Format

Alpha
|--Beta
`--Gamma
   |--Delta
   |*-Epsilon
   `--Zeta
      |?-Eta
      `--Theta 

Based roughly on Mike Keesey's old Dinosauricon site. Beta and Gamma are sister taxa within the clade Alpha Delta is within Gamma, but not a member of Zeta. Epsilon, however, is Gamma incertae sedis. Eta is only tentatively placed within Zeta. Nomina nuda are non-italicized and placed in quotation marks, while unpublished taxa (as recognized by the ICZN) are designated the same way, but followed by an asterisk. Nomina dubia are not distinguished from valid taxa, as which taxa are indeterminate is a subjective question. Taxa in parentheses are technically valid according to the ICZN but not used by most theropod workers.


Cladogram


Theropoda
|--Eodromaeus murphi `--+--Daemonosaurus chauliodus `--+--Tawa hallae
`--Avepoda
Averostra Eutheropoda |*-"Megalosaurus" woodwardi |*-"Newtonsaurus" cambrensis
|--Coelophysoidea (Podokesauroidea) | |?-Sarcosaurus woodi | |*-Halticosaurus longotarsus | |*-Sarcosaurus? andrewsi | `--+?-Dilophosauridae
| | |?-"Dilophosaurus" sinensis | | |?-Sinosaurus triassicus | | |?-Dracovenator regenti | | |?-Cryolophosaurus ellioti | | `--Dilophosaurus wetherilli | `--+*?"Comanchesaurus kuesi" | |*-Dolichosuchus cristatus | |*-Gojirasaurus quayi (in part) | |?-Procompsognathus triassicus
| |*-Protoavis texensis (in part) | |--Liliensternus liliensterni | `--+--Lophostropheus airelensis | |?-Zupaysaurus rougieri | `--Coelophysidae (Podokesauridae) | |*-Coelophysis? longicollis | |*?Coelophysis? willistoni | |*-Kayentavenator elysiae | |*-Longosaurus longicollis | |*-Podokesaurus holyokensis | |*-Pterospondylus trielbae | |*-"Syntarsus" "mexicanum" | |--Segisaurinae | | |--Camposaurus arizonensis | | |?-"Syntarsus" kayentakatae | | `--Segisaurus halli
| `--Coelophysinae
| |--Coelophysis bauri | `?-Coelophysis? rhodesiensis
`--Neotheropoda
|*-"Allosaurus" medius |*-Allosaurus? trihedrodon |*-Altispinax dunkeri |*-Calamospondylus oweni |*-"Capitalsaurus" potens |*-Chienkosaurus ceratosauroides |*-"Coelurosaurus" |*-Diplotomodon horrificus |*-"Elaphrosaurus" iguidiensis |*-"Futabasaurus" |*-"Katsuyamasaurus" |*?Kelmayisaurus? "gigantus" |*?"Labrosaurus" "huene" |*-"Megalosaurus" insignis |*-"Megalosaurus" lonzeensis |*-Megalosaurus? monasterii |*-"Megalosaurus" pombali |*?Megalosaurus? "tibetensis" |*?"Morosaurus" marchei |*-"Ornithocheirus" hilsensis |*-Orthogoniosaurus matleyi |*?Orthogoniosaurus? rawesi |*-Ozraptor subotaii |*-"Prodeinodon" kwangshiensis |*-Prodeinodon mongoliensis |*-"Scrotum humanum" |*-Szechuanosaurus campi |*-Teinurosaurus sauvagei |*-Wakinosaurus satoi |*-Walgettosuchus woodwardi |*?Yangchuanosaurus? "longqiaoensis"
|--Ceratosauria | |*?Bahariasaurus ingens | |*-"Merosaurus newmani" | |*?"Ngexisaurus dapukaensis" | |*-Xenotarsosaurus bonapartei | |--Berberosaurus liassicus | |--"Camarillasaurus cirugedae" | |--+--Deltadromeus agilis | | |?-Austrocheirus isasii | | |?-Ligabueino andesi | | |--Spinostropheus gautieri | | `--+--Elaphrosaurus bambergi | | `--Limusaurus inextricabilis | `--Neoceratosauria
| |--Ceratosauridae
| | |*?Ceratosaurus? roechlingi | | |--Genyodectes serus | | |?-+--Ostafrikasaurus crassiserratus | | | `--Ostafrikasaurus stechowi | | `--+--Ceratosaurus dentisulcatus | | |--Ceratosaurus magnicornis | | |--Ceratosaurus meriani | | |--Ceratosaurus nasicornis | | `--Ceratosaurus sulcatus | `--Abelisauroidea | |--Eoabelisaurus mefi | `--Abelisauria | |?-"Bayosaurus pubica" | |*?Betasuchus bredai | |*-"Nototyrannus violantei" | |*?Tarascosaurus salluvicus | |--Rahiolisaurus gujaratensis | |--Noasauridae | | |--Dahalokely tokana | | |*-Coeluroides largus | | |*-Compsosuchus solus | | |*-Jubbulpuria tenuis | | |*-Ornithomimoides? barasimlensis | | |*-Ornithomimoides mobilis | | |*-"Sidormimus"* | | |*-Velocisaurus unicus | | |*-Vitakrisaurus saraiki | | `--+--Genusaurus sisteronis | | |--Laevisuchus indicus | | |--Masiakasaurus knopfleri | | `--Noasaurus leali | `--+--Rugops primus | |*?"Carnosaurus" | |*-Dryptosauroides grandis | |*-Kryptops palaios | |*?Quilmesaurus curriei | `--Abelisauridae | |--"Majungasaurinae" | | |?-Arcovenator escotae | | |--Indosaurus matleyi | | |--Indosuchus raptorius | | |--Majungasaurus crenatissimus | | `--Rajasaurus narmadensis | `--Brachyrostra | |*-Pycnonemosaurus nevesi | |--Carnotaurinae | | Carnotaurini | | |--Aucasaurus garridoi | | `--Carnotaurus sastrei
| `--Abelisaurinae | |--Ekrixinatosaurus novasi | `--+--Skorpiovenator bustingorryi | `--+--Ilokelesia aguadagradensis | `--Abelisaurus comahuensis
`--Tetanurae |--Chuandongocoelurus primitivus |?-Monolophosaurus jiangi |*-Allosaurus? tendagurensis |*-Kaijiangosaurus lini |*?Megalosaurus? "dapukaensis" |*-"Megalosaurus" inexpectatus |*-"Megalosaurus" pannoniensis |*?"Mifunesaurus" |*?Prodeinodon? "tibetensis" |*-"Saltriosaurus" `-----Orionides |*-Cruxicheiros newmanorum |*-Dandakosaurus indicus |*-Ichthyovenator laosensis |*?"Megalosaurus" hungaricus |*-"Metriacanthosaurus" "reynoldsi" |*-Sigilmassasaurus brevicollis |--Megalosauroidea | |--Piatnitzkysauridae | | |--Marshosaurus bicentissimus | | `--+--Condorraptor currumili | | `--Piatnitzkysaurus floresi | `--Megalosauria | |*-Streptospondylus altdorfensis | |--Megalosauridae | | |*?Streptospondylus? cuvieri | | |--Eustreptospondylinae | | | `--Eustreptospondylus oxoniensis | | `--+--Megalosaurinae | | | |?-Duriavenator hesperis | | | `--+--Megalosaurus bucklandii | | | |*-"Megalosaurus" "phillipsi" | | | |*-"Brontoraptor" | | | |*-Edmarka rex | | | `--+--Torvosaurus tanneri | | | `--Torvosaurus gurneyi | | `--Afrovenatorinae | | |--Afrovenator abakensis | | |--Dubreuillosaurus valesdunensis | | |--Leshansaurus qianweiensis | | |?-Magnosaurus nethercombensis | | |--Piveteausaurus divesensis | | `?-Poekilopleuron bucklandii | `--Spinosauridae | |--Baryonychinae | | |--Baryonyx walkeri | | |--Suchosaurus cultridens | | |--Suchosaurus? girardi | | `--Cristatusaurus lapparenti | `--Spinosaurinae | |--Irritator challengeri | |--Oxalaia quilombensis | |--Siamosaurus suteethorni | |--Siamosaurus? fusuiensis | `--Spinosaurus aegyptiacus
`--Avetheropoda |*-"Tyrannosaurus" "lanpingensis" |*-Unquillosaurus ceibalei |*-Valdoraptor oweni
|--Carnosauria | Allosauroidea | |*-Erectopus superbus | |*?Kelmayisaurus petrolicus | |*-Xuanhanosaurus qilixiaensis | |--Metriacanthosauridae | | |--+--Yangchuanosaurus shangyouensis | | | `--Yangchuanosaurus zigongensis | | `--Metriacanthosaurinae | | |--Shidaisaurus jinae | | `--+--Metriacanthosaurus parkeri | | |--Siamotyrannus isanensis | | |--Sinraptor dongi | | `--Sinraptor? hepingensis | `--Allosauria | |--Carcharodontosauridae | | |--Neovenator salerii | | `--+--+--Becklespinax altispinax | | | |--Concavenator corcovatus | | | |--Eocarcharia dinops | | | `--Sauroniops pachytholus | | `--+--+--Acrocanthosaurus atokensis | | | `--Veterupristisaurus milneri
| | `--+--+?-Labocania anomala | | | `--Shaochilong maortuensis | | `--Carcharodontosaurinae | | |*-"Megalosaurus" chubutensis | | |*?"Megalosaurus" ingens | | |--+--Carcharodontosaurus saharicus | | | `--Carcharodontosaurus? iguidensis | | `--Giganotosaurinae | | |--Tyrannotitan chubutensis | | `--+--Mapusaurus roseae | | `--Giganotosaurus carolinii | `--Allosauridae | |*-Antrodemus valens | |*?"Madsenius trux" | |*-Saurophaganax maximus | |*-"Wyomingraptor" | `--+--Allosaurus "jimmadseni" | `--+--Allosaurus atrox | |--Allosaurus fragilis | `--Allosaurus europaeus
`--Coelurosauria |?-Bicentenaria argentina |?-Gasosaurus constructus |?-Lourinhanosaurus antunesi |?-Sciurumimus albersdoerferi |?-Tugulusaurus faciles |*-Iliosuchus incognitus |*-Zuolong salleei `--+*-Xinjiangovenator parvus |?-Megaraptora | |*-Rapator ornitholestoides | |?-Chilantaisaurus tashuikouensis | `--+*-"Allosaurus" sibiricus | |--Siats meekerorum | |--+--"Allosaurus" "robustus" | | |--Australovenator wintonensis | | `--Fukuiraptor kitadaniensis | `--+--Megaraptor namunhuaiquii | |--Aerosteon riocoloradensis | `--Orkoraptor burkei `--Tyrannoraptora |*-Aristosuchus pusillus |*-"Arkansaurus fridayi" |*-"Dryosaurus" grandis |*?Kakuru kujani
|*?Microvenator? "chagyabi" |*-Santanaraptor placidus |*-Timimus hermani |*-"Tonouchisaurus mongoliensis" |*-Yutyrannus huali |--Tyrannosauroidea (Deinodontoidea) | |*-Aviatyrannis jurassica | |--Alectrosaurus olseni | `--+--+--Stokesosaurus clevelandi | | `--Juratyrant langhami | `--+?-Embasaurus minax | |--Xiongguanlong baimoensis | `--+--+--Dryptosaurus aquilunguis
| | `--Dryptosaurus? macropus | `--+--Raptorex kriegsteini | `--+--Appalachiosaurus montgomeriensis | `--+--Bistahieversor sealeyi | `--Tyrannosauridae (Deinodontidae) | |*-Aublysodon? lateralis | |*-Deinodon horridus | |*-Deinodon? falculus | |*?Deinodon? grandis | |*-Deinodon? hazenianus | |*-"Ornithomimus" tenuis | |*?Chingkankousaurus fragilis | |*-Tarbosaurus? periculosus | |--Albertosaurinae | | |--Gorgosaurus libratus | | |--"Albertosaurus" incrassatus | | `--Albertosaurus sarcophagus | `--Tyrannosaurinae | |--Teratophoneus curriei | `--+*-Aublysodon mirandus | |--Daspletosaurus torosus | `--+--Lythronax argestes | `--+--Nanuqsaurus hoglundi | `--+--+--Alioramus remotus | | `--Tarbosaurus bataar | |--Zhuchengtyrannus magnus | |?-Tyrannosaurus? zhuchengensis | `--Tyrannosaurus rex `--Avipluma |--Coelurus fragilis |--Eotyrannus lengi |--Tanycolagreus topwilsoni |--+--Calamosaurus foxii | |--Dilong paradoxus | `?-Nuthetes destructor `--+*-"Beelemodon" |*-"Kagasaurus" |*-Phaedrolosaurus ilikensis |--Proceratosauridae | |?-Aorun zhaoi
| |--Guanlong wucaii | |--Kileskus aristotocus | |--Sinotyrannus kazuoensis | `--Proceratosaurus bradleyi |--Mirischia asymmetrica |--Sinocalliopteryx gigas `--+--Nedcolbertia justinhofmanni `--+?-Bagaraatan ostromi |--Compsognathidae | |--Huaxiagnathus orientalis | `--+--Sinosauropteryx prima | `--Compsognathus longipes |?-Juravenator starki `--+--Ornitholestes hermanni |--Scipionyx samniticus `-----Avepectora
Metornithes |*-Yaverlandia bitholus |--Arctometatarsalia | |*-Kinnareemimus khonkaenensis | Ornithomimoidea | |--Ornithomimosauria | | |*-"Sanchusaurus" | | |--Hexing qingyi | | `--+--Deinocheirus mirificus | | `--+--Pelecanimimus polyodon | | `--+--Harpymimidae | | | |--Harpymimus okladnikovi | | | `--Shenzhousaurus orientalis | | `--Ornithomiminae | | |--Beishanlong grandis | | `--+--Garudimimus brevipes | | `--Ornithomimidae | | |--"Grusimimus tsuru"* | | `--+--Sinornithomimus dongi | | |--Qiupalong henanensis | | |*-Archaeornithomimus? bissektensis | | `--+*-"Coelosaurus" antiquus | | |--+--Anserimimus planinychus | | | `--+--Dromiceiomimus brevitertius | | | `--Dromiceiomimus samueli | | `--+--Struthiomimus altus | | `--+--Gallimimus bullatus | | |*-"Orcomimus" | | `--+--Ornithomimus velox | | `--+--Ornithomimus? sedens | | `--+--Archaeornithomimus asiaticus | | `--"Gallimimus" "mongoliensis" | `--Alvarezsauroidea | |*-Alnashetri cerropoliciensis | |?-Nqwebasaurus thwazi | |--Aniksosaurus darwini | `--+--Haplocheirus sollers | `--Alvarezsauridae | |--+--Achillesaurus manazzonei | | `--Alvarezsaurus calvoi | `--+--Patagonykinae | | |--Patagonykus puertai | | `--Bonapartenykus ultimus | `--+*?Kol ghuva | |--Linhenykus monodactylus | `--+--Xixianykus zhangi | `--Parvicursorinae | |*-+--Albertonykus borealis | | `--"Ornithomimus" minutus | |--Parvicursor remotus | `--Mononykini | |*-Heptasteornis andrewsi | |--+--Albinykus baatar | | `--Shuvuuia deserti | `--+--Ceratonykus oculatus | `--Mononykus olecranus `--Maniraptora Chuniaoae |*-Bradycneme draculae |*-Ilerdopteryx viai |*-Martharaptor greenriverensis |*-Ningyuansaurus wangi |*?Praeornis sharovi |*-Shanyangosaurus niupanggouensis |--Enigmosauria | |--Therizinosauria
| | |?*Thecocoelurus daviesi
| | |--Falcarius utahensis | | `--+?-Eshanosaurus deguchiianus | | |--Jianchangosaurus yixianensis
| | `--Therizinosauroidea
| | |*-"Chilantaisaurus" zheziangensis
| | |*?Inosaurus tedreftensis
| | |*-"Nanshiungosaurus" bohlini | | |--Beipiaosaurus inexpectus
| | `--+--Alxasaurus elesitaiensis | | `--+--Erliansaurus bellamanus | | `--+--Neimongosaurus yangi | | `--Therizinosauridae | | |*-Erlikosaurus andrewsi
| | |*-Therizinosaurus cheloniformis | | |--Enigmosaurus mongoliensis | | `--+--Suzhousaurus megatherioides | | `--+--+--Nothronychus mckinleyi | | | `--Nothronychus? graffami | | `--+--Nanshiungosaurus brevispinus | | `--Segnosaurus galbinensis | `--Oviraptorosauria
| |--+--Incisivosaurus gauthieri | | `--Protarchaeopteryx robusta | `--+--Similicaudipteryx yixianensis | `--+*-Luoyanggia liudianensis | |--Caudipteryx zoui | `--Caenagnathoidea | |*-Ganzhousaurus nankangensis | |--Caenagnathidae
| | |*-Caenagnathasia martinsoni | | |*-Hagryphus giganteus | | |*-Kuszholia mengi | | |*-Ojoraptorsaurus boerei | | |?-Nomingia gobiensis | | |--Chirostenotes pergracilis | | |--Anzu wyliei | | `--Elmisaurinae | | |*-Shixinggia oblita | | |--Avimimus portentosus | | `--+--Elmisaurus rarus | | |--Elmisaurus? elegans | | `--Leptorhynchos gaddisi | `--Oviraptoridae | |?-Gigantoraptor erlianensis | `--+--Nankangia jiangxiensis | `--+?-Yulong mini | `--Oviraptorinae
| |?-Microvenator celer | |--Banji long | |--Oviraptor philoceratops | |--Wulatelong gobiensis | `--+--Citipati osmolskae | |--Rinchenia mongoliensis | `--+*-Jiangxisaurus ganzhouensis | |--Khaan mckannai | `--"Ingeniinae" | |--Conchoraptor gracilis | `--+--Machairasaurus leptonychus | `--+--"Ingenia" yanshini | `--+--Heyuannia huangi | `--Nemegtomaia barsboldi `--Paraves `--Eumaniraptora |*-Anchiornis huxleyi |*-Archaeornithoides deinosauriscus |*-Aurornis xui |*-Austroraptor cabazai |*-Balaur bondoc |*-Bauxitornis mindszentyae
|*-Buitreraptor gonzalezorum |*-Cretaaviculus sarysuensis |*-Elopteryx nopcsai |*-Eosinopteryx brevipenna |*-Mahakala omnogovae |*-Neuquenraptor argentinus |*-Pamparaptor micros |*-Pedopenna daohugouensis |*-Pneumatoraptor fodori |*-Rahonavis ostromi |--Unenlagiinae | |--Unenlagia comahuensis | `--Unenlagia? paynemili |*-Wyleyia valdensis |*-Xiaotingia zhengi |*-Yixianosaurus longimanus |--Deinonychosauria
| Dromaeosauridae (Ornithodesmidae) | `--+--Microraptoria | | |?-Bambiraptor feinbergi | | `--+--Graciliraptor lujiatunensis | | |--"Paleopteryx thomsoni" | | |--Sinornithosaurus millenii | | |?-+--Richardoestesia gilmorei | | | `?-+--Richardoestesia? isosceles | | | `--Richardoestesia? asiatica | | |--Microraptor zhaoianus | | |--Hesperonychus elizabethae | | `?-Shanag ashile | |*-Itemirus medullaris | |*-"Kitadanisaurus" | |*-Tianyuraptor ostromi | `--Eudromaeosauria | |--Saurornitholestinae | | |--Saurornitholestes langstoni | | `--Saurornitholestes? robustus | |--Velociraptorinae
| | |--Velociraptor mongoliensis | | |--Velociraptor osmolskae | | |--Adasaurus mongoliensis | | `?-Luanchuanraptor henanensis | |--Deinonychus antirrhopus | |--Tsaagan mangas | |--Acheroraptor temertyorum | |*-"Dromaeosaurus" gracilis | |*-"Ichabodcraniosaurus" | |*-"Koreanosaurus koreanensis" | |*-Ornithodesmus cluniculus | |*-Pyroraptor olympius | |*-Variraptor mechinorum | `--Dromaeosaurinae | |*-Dromaeosauroides bornholmensis | |*-Yurgovuchia doellingi | |*-Zapsalis abradens | |--+?-Atrociraptor marshalli | | `--Dromaeosaurus albertensis | `?-+--Achillobator giganticus | `--Utahraptor ostrommaysorum |--Troodontidae | |*?+--Paronychodon lacustris | | |--Paronychodon caperatus | | |--Paronychodon asiaticus | | `--Paronychodon portucalensis | |*-Urbacodon itemirensis | |*-Xixiasaurus henanensis | |--Mei long | `--+--Sinovenator changii | |?-Hulsanpes perlei | |?-Jinfengopteryx elegans | `--+--Sinusonasus magnodens | |--Koparion douglassi | `--+--Byronosaurus jaffei | |--Geminiraptor suarezarum | |--Gobivenator mongoliensis | `--+*-"Saurornithoides" asiamericanus | |--Sinornithoides youngi | `--Troodontinae | |*-Borogovia gracilicrus | |*-Tochisaurus nemegtensis | |--Saurornithoides mongoliensis | `--+--Zanabazar junior | |--Linhevenator tani | |?-Philovenator curriei | |--Talos sampsoni | |--Troodon formosus | `--Troodon bakkeri `--Avialae |?-Scansoriopterygidae | |--Epidexipteryx hui | `--Scansoriopteryx heilmanni `-----Ornithes |?-Archaeopterygidae | `--Archaeopteryx lithographica `--Ornithurae |*?Cerebavis cenomanica |*-Dalianraptor cuhe |--Shenzhouraptor sinensis `--Euavialae |--Jixiangornis orientalis |--Yandangornis longicaudus |--Zhongjianornis yangi `--Avebrevicauda |--Omnivoropterygiformes | `--Sapeornis chaoyangensis `--------Pygostylia |*-Qiliania graffini |--Confuciusornithiformes | Confuciusornithidae | |*-"Proornis coreae" | |--Changchengornis hengdaoziensis | `--+*-Zhongornis haoae | |*-Confuciusornis chuonzhous | |--Confuciusornis dui | |--Confuciusornis feducciai | |--Confuciusornis zhengi | `--Confuciusornis sanctus `--Ornithothoraces |*-Aberratiodontus wui |*-"Confuciusornis" jianchangensis |*-Hollanda luceria |*-Lectavis bretincola |*?Lenesornis maltshevskyi |*-Platanavis nana |*-Xinghaiornis lini |--Enantiornithes | |*-Abavornis bonaparti | |*-Catenoleimus anachoretus | |*-Cuspirostrisornis houi | |*-Enantiophoenix electrophyla | |*-"Enantiornis" walkeri | |*-Gracilornis jiufotangensis | |*-"Holbotia ponomarenkoi" | |*-Huoshanornis huji | |*-Iberomesornis romerali | |*-"Ichthyornis" minusculus | |*-+--Incolornis silvae | | `--Incolornis martini | |*-Longchengornis sanyanensis | |--Explorornis nessovi | |--Noguerornis gonzalezi | |--Pengornis houi | |--Protopteryx fengningensis | `--+*-Alexornis antecedens | |*-Halimornis thompsoni | |*-Kizylkumavis cretacea | |*-Sazavis prisca | |--Boluochia zhengi | |--Longipteryx chaoyangensis | |--Longipteryx? yangi | |--Shengjingornis yangi | |--Longirostravisidae | | |--Shanweiniao cooperorum | | `--+--Longirostravis hani | | `--Rapaxavis pani | `--+*-Alethoalaornis agitornis | |*-Gurilynia nessovi | |--+--Avisaurus archibaldi | | `--Avisaurus gloriae | |--Cathayornis? aberransis | |--Cathayornis? caudatus | |--Cathayornis? chabuensis | |--Cathayornis yandica | |--Concornis lacustris | |--Dalingheornis liweii | |--Dapingfangornis sentisorhinus | |--Elbretornis bonapartei | |--Elsornis keni | |--Enantiornis leali | |--Eocathayornis walkeri | |--Eoenantiornis buhleri | |--Flexomornis howei | |--Gobipipus reshetovi | |--Gobipteryx minuta | |--Hebeiornis fengningensis | |--Intiornis inexpectatus | |--Jibeinia luanhera | |--Largirostrornis sexdentoris | |--Liaoxiornis delicatus | |--Martinavis cruzyensis | |--Martinavis? minor | |--Martinavis? saltariensis | |--Martinavis? vincei | |--Martinavis? whetstonei | |--Mystiornis cyrili | |--Nanantius eos | |--Neuquenornis volans | |--Otogornis genghisi | |--Paraprotopteryx gracilis | |--Parvavis chuxiongensis | |--Sinornis santensis | |--Soroavisaurus australis | |--Xiangornis shenmi | |--Yungavolucris brevipedalis | |--Bohaiornithidae | | |--+--Shenqiornis mengi | | | `--Sulcavis geeorum | | `--+--Zhouornis hani | | `--+--Longusunguis kurochkini | | `--+--Parabohaiornis martini | | `--Bohaiornis guoi | `--Liaoningornithidae | |?-Vorona berivotrensis | |--Eoalulavis hoyasi | `--Liaoningornis longidigitus
`--+*-Jiuquanornis niui |*-Parahongshanornis chaoyangensis |--Archaeorhynchus spathula |--Chaoyangiformes | `--Chaoyangia beishanensis `--Ornithuromorpha |*-Gargantuavis philoinos |*-"Zhyraornis kashkarovi" |--Hongshanornis longicresta |--Jianchangornis microdonta |--Piscivoravis lii |--Schizooura lii |--Yumenornis huangi |--Patagopterygiformes | |--Patagopteryx deferrariisi | `?-Alamitornis minutus `--+--Longicrusavis houi `--+--Songlingornithidae | |--+--Yanornis martini | | `--Yanornis guozhangi | `--+--Yixianornis grabaui | `--Songlingornis linghensis `--+*-Changmaornis houi |*-Guildavis tener |*-+--Zhyraornis kashkarovi | `--Zhyraornis logunovi |--Gansuiformes | `--Gansus yumenensis `--+*-Horezmavis eocretacea |--Ichthyornithes | `--Ichthyornis anceps `--+--Hesperornithes | Hesperornithiformes | |*-Potamornis skutchi | |--Enaliornis barretti | |--Enaliornis? sedgwicki | |--Enaliornis? seeleyi | |--Pasquiaornis hardiei | |--Pasquiaornis? tankei | `--Hesperornithoidea | |*-Hesperornis? macdonaldi | |*-Judinornis nogontsavensis | |*-Parascaniornis stensioei | |--Baptornithidae | | `--Baptornis advenus | `--Hesperornithidae | |--+?-Brodavis americanus | | |?-Brodavis? baileyi | | |--Brodavis? varneri | | `?-Brodavis? mongoliensis | `--+--Parahesperornis alexi | |?-Hesperornis? mengeli | `--+--Hesperornis bairdi | `--+*?Canadaga arctica | |*-Hesperornis altus | |*-Hesperornis montanus | |--Hesperornis gracilis | |--Hesperornis chowi | |--Hesperornis regalis | `--+--Hesperornis crassipes | |--Hesperornis rossicus | `--Hesperornis bazhanovi `--Carinatae |--Ambiortiformes | |--Ambiortus dementjevi | |--Apsaravis ukhaana | `--Palintropus retusus `-----Aves |*-Apatornis celer |*-Ceramornis major |*-Gallornis straeleni |*-Iaceornis marshi |*-Laornis edvardsianus |*-+--Palaeotringa littoralis | `--Palaeotringa vagans |*-"Palaeotringa" vetus
|*-Volgavis marina |--Palaeognathae | |?-Limenavis patagonica | |--Struthionidae | `--Notopalaeognathae | |--Rheidae | `--+--+--Dinornithidae | | `--Tinamidae | `--Novaeratitae | |--Apterygidae | `--Casuariidae `--Neognathae |*-Qinornis paleocenica |--Galloanserae | |--Pangalliformes | | |*-Austinornis lentus | | |--Quercymegapodiidae | | `--Galliformes | | |--Megapodiidae | | `--+--Cracidae | | `--+--Numididae | | `--Phasianidae | `--Anseriformes | |--Anhimidae | `--Anseres | |--Anseranatidae | `--+--Anatalavis rex | `--Anatoidea | |--Vegavis iaai | |--Presbyornithidae | | `--Teviornis gobiensis | `--Anatidae `--Neoaves |--Opisthocomidae |--+--+--Lithoptila | | `--+--Prophaethon | | `--Phaethontidae | `--Eurypygoidea | |--Eurypygidae | `--Rhynochetidae |--Pteroclididae |--Columbidae |--Mesitornithidae |--Mirandornithes | |--Podicipedidae | `--Phoenicopteridae |--Cypselomorphae | |--Steatornithidae | |--Nyctibiidae | |--Podargidae | |--Caprimulgidae | `--Daedalornithes | |--Aegothelidae | `--Apodiformes | |--Trochillidae | `--Apodi | |--Hemiprocnidae | `--Apodidae `--Coronaves |--Insolitaves | |--Otididae | |--Cuculidae | |--Musophagidae | |--Gruiformes | | |--Ralloidea | | | |--Heliornithidae | | | `--Rallidae | | `--Grues | | |--Psophiidae | | `--Gruoidea | | |--Aramidae | | `--Gruidae | `--Natatores | |--Gaviiformes | | |--Neogaeornis wetzeli | | `--Polarornis gregorii | |--Austrodyptornithes | | |--Sphenisciformes | | `--Procellariiformes | | |?-Lonchodytes estesi | | |--Tytthostonyx glauconiticus | | |--Diomedeidae | | `--+--Oceanitidae | | `--Procellariidae | `--Steganograllatores | |--Ciconiidae | `--Pelecaniformes | |*?Torotigidae | | |?-Novacaesareala hungerfordi | | `--Torotix clemensi | |--Steganopodes | | |--Fregatidae | | `--Sulida | | |--Sulidae | | `--Phalacrocoracoidea | | |--Anhingidae | | `--Phalacrocoracidae | `--+--Ardeidae | |--Threskiornithidae | `--+--Scopidae | |--Balaenicipitidae | `--Pelecanidae `--Litoritelluraves |--Charadriiformes | |*?Cimolopteryx rara | |*?"Cimolopteryx" maxima | |*?"Cimolopteryx" minima | |*?"Cimolopteryx" petra | |?-+--Graculavus velox | | `--Graculavus augustus | |?-Telmatornis priscus | |?-"Lonchodytes" pterygius | |--Charadrii | | |--+--Charadriidae | | | `--Haematopoidea | | | |--Recurvirostridae | | | `--Haematopodidae | | `--Chionidi | | |--Burhinidae | | `--+--Chionididae | | `--Pluvianellidae | `--+--Scolopaci | | |--Scolopacidae | | `--Thinocori | | |--Thinocoroidea | | | |--Pedionomidae | | | `--Thinocoridae | | `--Parrae | | |--Rostratulidae | | `--Jacanidae | `--Lari | |--Turnicidae | `--+--Glareolidae | `--+--Dromadidae | `--+--Alcidae | |--Stercorariidae | `--Larides | |--Laridae | |--Sternidae | `--Rhynchopidae `--Dendrornithes |--Accipitriformes | |--Cathartidae | `--Accipitres | |--Sagittariidae | `--+--Pandionidae | `--Accipitridae |--+--+--Coliidae | | `--Strigiformes | | |--Tytonidae | | `--Strigidae | --Afroaves | |--Leptosomidae | `--Eucavitaves | |--Trogonidae | `--Picocoraciae | |--Bucerotiformes | | |--Bucerotidae | | `--Upupides | | |--Phoeniculidae | | `--Upupidae | `--Picodynastornithes | |--Coraciiformes | | |--Meropidae | | |--Coracioidea | | | |--Brachypteraciidae | | | `--Coraciidae | | `--Halcyones | | |--Todidae | | `--Alcedinoidea | | |--Motmotidae | | `--Alcedinidae | `--Piciformes | |--Galbulae | | |--Bucconidae | | `--Galbulidae | `--Pici | |--Capitonoidea | | |--Capitonidae | | `--Rhamphastidae | `--Picoidea | |--Indicatoridae | `--Picidae `--Australavis |--Cariamidae `--Eufalconimorphae |--Falconidae `--Psittacopasserae |--Psittaciformes | |--Nestoridae | `--+--Cacatuidae | `--Psittacidae `--Passeriformes |--Acanthisittidae `--Eupasseres |--Tyranni | |--Eurylaimides | `--Tyrannides | |--Tyrannoidea | `--Furnarioidea `--Passeri |--Menuridae `--+--Climacteridae `--+--Meliphagoidea `--+--Corvoidea `--+--Picathartidae `--Passerida